Author Correction: Genetic meta-analysis of diagnosed Alzheimer's disease identifies new risk loci and implicates Aβ, tau, immunity and lipid processing

Brian W Kunkle^#¹, Benjamin Grenier-Boley^#^{2

3

4}, Rebecca Sims^{5

6}, Joshua C Bis⁷, Vincent Damotte^{2

3

4}, Adam C Naj⁸, Anne Boland⁹, Maria Vronskaya⁵, Sven J van der Lee¹⁰, Alexandre Amlie-Wolf¹¹, Céline Bellenguez^{2

3

4}, Aura Frizatti⁵, Vincent Chouraki^{2

3

4

12

13}, Eden R Martin¹⁴, Kristel Sleegers^{15

16}, Nandini Badarinarayan⁵, Johanna Jakobsdottir¹⁷, Kara L Hamilton-Nelson¹⁴, Sonia Moreno-Grau^{18

19}, Robert Olaso⁹, Rachel Raybould^{5

6}, Yuning Chen²⁰, Amanda B Kuzma¹¹, Mikko Hiltunen^{21

22}, Taniesha Morgan⁵, Shahzad Ahmad¹⁰, Badri N Vardarajan^{23

24

25}, Jacques Epelbaum²⁶, Per Hoffmann^{27

28

29}, Merce Boada^{18

19}, Gary W Beecham¹⁴, Jean-Guillaume Garnier⁹, Denise Harold³⁰, Annette L Fitzpatrick^{31

32}, Otto Valladares¹¹, Marie-Laure Moutet⁹, Amy Gerrish³³, Albert V Smith^{34

35}, Liming Qu¹¹, Delphine Bacq⁹, Nicola Denning^{5

6}, Xueqiu Jian³⁶, Yi Zhao¹¹, Maria Del Zompo³⁷, Nick C Fox^{33

38}, Seung-Hoan Choi¹⁸, Ignacio Mateo³⁹, Joseph T Hughes⁴⁰, Hieab H Adams¹⁰, John Malamon¹¹, Florentino Sanchez-Garcia⁴¹, Yogen Patel⁴⁰, Jennifer A Brody⁷, Beth A Dombroski¹¹, Maria Candida Deniz Naranjo⁴¹, Makrina Daniilidou⁴², Gudny Eiriksdottir¹⁷, Shubhabrata Mukherjee⁴³, David Wallon⁴⁴, James Uphill⁴⁵, Thor Aspelund^{17

46}, Laura B Cantwell¹¹, Fabienne Garzia⁹, Daniela Galimberti^{47

48}, Edith Hofer^{49

50}, Mariusz Butkiewicz⁵¹, Bertrand Fin⁹, Elio Scarpini^{47

48}, Chloe Sarnowski²⁰, Will S Bush⁵¹, Stéphane Meslage⁹, Johannes Kornhuber⁵², Charles C White⁵³, Yuenjoo Song⁵¹, Robert C Barber⁵⁴, Sebastiaan Engelborghs^{55

56}, Sabrina Sordon⁵⁷, Dina Voijnovic¹⁰, Perrie M Adams⁵⁸, Rik Vandenberghe⁵⁹, Manuel Mayhaus⁵⁷, L Adrienne Cupples^{12

20}, Marilyn S Albert⁶⁰, Peter P De Deyn^{55

56}, Wei Gu⁵⁷, Jayanadra J Himali^{12

13

20}, Duane Beekly⁶¹, Alessio Squassina³⁷, Annette M Hartmann⁶², Adelina Orellana¹⁸, Deborah Blacker^{63

64}, Eloy Rodriguez-Rodriguez³⁹, Simon Lovestone⁶⁵, Melissa E Garcia⁶⁶, Rachelle S Doody⁶⁷, Carmen Munoz-Fernadez⁴¹, Rebecca Sussams⁶⁸, Honghuang Lin⁶⁹, Thomas J Fairchild⁷⁰, Yolanda A Benito⁴¹, Clive Holmes⁶⁸, Hata Karamujić-Čomić¹⁰, Matthew P Frosch⁷¹, Hakan Thonberg^{72

73}, Wolfgang Maier^{74

75}, Gennady Roshchupkin¹⁰, Bernardino Ghetti⁷⁶, Vilmantas Giedraitis⁷⁷, Amit Kawalia⁷⁸, Shuo Li²⁰, Ryan M Huebinger⁷⁹, Lena Kilander⁷⁷, Susanne Moebus⁸⁰, Isabel Hernández^{18

19}, M Ilyas Kamboh^{81

82

83}, RoseMarie Brundin⁷⁷, James Turton⁸⁴, Qiong Yang²⁰, Mindy J Katz⁸⁵, Letizia Concari^{86

87}, Jenny Lord⁸⁴, Alexa S Beiser^{12

13

20}, C Dirk Keene⁸⁸, Seppo Helisalmi^{21

22}, Iwona Kloszewska⁸⁹, Walter A Kukull³², Anne Maria Koivisto^{21

22}, Aoibhinn Lynch^{90

91}, Lluís Tarraga^{18

19}, Eric B Larson⁹², Annakaisa Haapasalo⁹³, Brian Lawlor^{90

91}, Thomas H Mosley⁹⁴, Richard B Lipton⁸⁵, Vincenzo Solfrizzi⁹⁵, Michael Gill^{90

91}, W T Longstreth Jr^{32

96}, Thomas J Montine⁸⁸, Vincenza Frisardi⁹⁷, Monica Diez-Fairen^{98

99}, Fernando Rivadeneira^{10

100

101}, Ronald C Petersen¹⁰², Vincent Deramecourt¹⁰³, Ignacio Alvarez^{98

99}, Francesca Salani¹⁰⁴, Antonio Ciaramella¹⁰⁴, Eric Boerwinkle^{105

106}, Eric M Reiman^{107

108

109

110}, Nathalie Fievet^{2

3

4}, Jerome I Rotter¹¹¹, Joan S Reisch¹¹², Olivier Hanon¹¹³, Chiara Cupidi¹¹⁴, A G Andre Uitterlinden^{10

100

101}, Donald R Royall¹¹⁵, Carole Dufouil^{116

117}, Raffaele Giovanni Maletta¹¹⁴, Itziar de Rojas^{18

19}, Mary Sano¹¹⁸, Alexis Brice^{119

120}, Roberta Cecchetti¹²¹, Peter St George-Hyslop^{122

123}, Karen Ritchie^{124

125

126}, Magda Tsolaki⁴², Debby W Tsuang^{127

128}, Bruno Dubois^{129

130

131

132}, David Craig¹³³, Chuang-Kuo Wu¹³⁴, Hilkka Soininen^{21

22}, Despoina Avramidou⁴², Roger L Albin^{135

136

137}, Laura Fratiglioni¹³⁸, Antonia Germanou⁴², Liana G Apostolova^{139

140

141

142}, Lina Keller¹³⁸, Maria Koutroumani⁴², Steven E Arnold¹⁴³, Francesco Panza⁹⁷, Olymbia Gkatzima⁴², Sanjay Asthana^{144

145

146}, Didier Hannequin⁴⁰, Patrice Whitehead¹⁴, Craig S Atwood^{140

141

142}, Paolo Caffarra^{83

84}, Harald Hampel^{147

148

149

150}, Inés Quintela¹⁵¹, Ángel Carracedo¹⁵¹, Lars Lannfelt⁷⁷, David C Rubinsztein^{122

152}, Lisa L Barnes^{153

154

155}, Florence Pasquier¹⁰³, Lutz Frölich¹⁵⁶, Sandra Barral^{23

24

25}, Bernadette McGuinness¹³³, Thomas G Beach¹⁵⁷, Janet A Johnston¹³³, James T Becker^{81

158

159}, Peter Passmore¹³³, Eileen H Bigio^{160

161}, Jonathan M Schott³³, Thomas D Bird^{96

127}, Jason D Warren³³, Bradley F Boeve¹⁰², Michelle K Lupton^{40

162}, James D Bowen¹⁶³, Petra Proitsi⁴⁰, Adam Boxer¹⁶⁴, John F Powell⁴⁰, James R Burke¹⁶⁵, John S K Kauwe¹⁶⁶, Jeffrey M Burns¹⁶⁷, Michelangelo Mancuso¹⁶⁸, Joseph D Buxbaum^{118

169

170}, Ubaldo Bonuccelli¹⁶⁸, Nigel J Cairns¹⁷¹, Andrew McQuillin¹⁷², Chuanhai Cao¹⁷³, Gill Livingston¹⁷², Chris S Carlson^{145

146}, Nicholas J Bass¹⁷², Cynthia M Carlsson¹⁷⁴, John Hardy¹⁷⁵, Regina M Carney¹⁷⁶, Jose Bras^{38

177}, Minerva M Carrasquillo¹⁷⁸, Rita Guerreiro^{38

177}, Mariet Allen¹⁷⁸, Helena C Chui¹⁷⁹, Elizabeth Fisher¹⁷⁷, Carlo Masullo¹⁸⁰, Elizabeth A Crocco¹⁸¹, Charles DeCarli¹⁸², Gina Bisceglio¹⁷⁸, Malcolm Dick¹⁸³, Li Ma¹⁷⁸, Ranjan Duara¹⁸⁴, Neill R Graff-Radford¹⁷⁸, Denis A Evans¹⁸⁵, Angela Hodges¹⁸⁶, Kelley M Faber¹³⁹, Martin Scherer¹⁸⁷, Kenneth B Fallon¹⁸⁸, Matthias Riemenschneider⁵⁷, David W Fardo¹⁸⁹, Reinhard Heun⁷⁵, Martin R Farlow¹⁴¹, Heike Kölsch⁷⁵, Steven Ferris¹⁹⁰, Markus Leber¹⁹¹, Tatiana M Foroud¹³⁹, Isabella Heuser¹⁹², Douglas R Galasko¹⁹³, Ina Giegling⁶², Marla Gearing^{194

195}, Michael Hüll¹⁹⁶, Daniel H Geschwind¹⁹⁷, John R Gilbert¹⁴, John Morris^{198

199}, Robert C Green²⁰⁰, Kevin Mayo^{198

201

202}, John H Growdon²⁰³, Thomas Feulner⁵⁷, Ronald L Hamilton²⁰⁴, Lindy E Harrell²⁰⁵, Dmitriy Drichel²⁰⁶, Lawrence S Honig²³, Thomas D Cushion^{5

6}, Matthew J Huentelman¹⁰⁷, Paul Hollingworth⁵, Christine M Hulette²⁰⁷, Bradley T Hyman²⁰³, Rachel Marshall⁵, Gail P Jarvik^{208

209}, Alun Meggy⁵, Erin Abner²¹⁰, Georgina E Menzies^{5

6}, Lee-Way Jin²¹¹, Ganna Leonenko⁵, Luis M Real²¹¹, Gyungah R Jun²¹², Clinton T Baldwin²¹², Detelina Grozeva⁵, Anna Karydas¹⁶³, Giancarlo Russo²¹³, Jeffrey A Kaye^{214

215}, Ronald Kim²¹⁶, Frank Jessen^{74

75

191}, Neil W Kowall^{13

217}, Bruno Vellas²¹⁸, Joel H Kramer²¹⁹, Emma Vardy²²⁰, Frank M LaFerla²²¹, Karl-Heinz Jöckel⁸⁰, James J Lah²²², Martin Dichgans^{223

224}, James B Leverenz²²⁵, David Mann²²⁶, Allan I Levey²²², Stuart Pickering-Brown²²⁶, Andrew P Lieberman²²⁷, Norman Klopp²²⁸, Kathryn L Lunetta²⁰, H-Erich Wichmann^{229

230

231}, Constantine G Lyketsos²³², Kevin Morgan²³³, Daniel C Marson²⁰⁵, Kristelle Brown⁸⁴, Frank Martiniuk²³⁴, Christopher Medway⁸⁴, Deborah C Mash²³⁵, Markus M Nöthen^{27

28}, Eliezer Masliah^{193

236}, Nigel M Hooper²²⁶, Wayne C McCormick⁴³, Antonio Daniele²³⁷, Susan M McCurry²³⁸, Anthony Bayer²³⁹, Andrew N McDavid¹⁷³, John Gallacher⁶⁵, Ann C McKee^{13

217}, Hendrik van den Bussche¹⁸⁷, Marsel Mesulam^{161

240}, Carol Brayne²⁴¹, Bruce L Miller²⁴², Steffi Riedel-Heller²⁴³, Carol A Miller²⁴⁴, Joshua W Miller²⁴⁵, Ammar Al-Chalabi²⁴⁶, John C Morris^{171

201}, Christopher E Shaw^{246

247}, Amanda J Myers¹⁸¹, Jens Wiltfang^{248

249

250}, Sid O'Bryant⁵⁴, John M Olichney¹⁸², Victoria Alvarez²⁵¹, Joseph E Parisi²⁵², Andrew B Singleton²⁵³, Henry L Paulson^{135

137}, John Collinge⁴⁵, William R Perry¹⁴, Simon Mead⁴⁵, Elaine Peskind¹²⁸, David H Cribbs²⁵⁴, Martin Rossor³³, Aimee Pierce²⁵⁴, Natalie S Ryan⁴⁵, Wayne W Poon¹⁸³, Benedetta Nacmias^{255

256}, Huntington Potter²⁵⁷, Sandro Sorbi^{255

258}, Joseph F Quinn^{187

188}, Eleonora Sacchinelli¹⁰⁴, Ashok Raj¹⁷³, Gianfranco Spalletta^{259

260}, Murray Raskind¹²⁸, Carlo Caltagirone²⁵⁹, Paola Bossù¹⁰⁴, Maria Donata Orfei²⁵⁹, Barry Reisberg^{190

261}, Robert Clarke²⁶², Christiane Reitz^{23

24

263}, A David Smith²⁶⁴, John M Ringman²⁶⁵, Donald Warden²⁶⁴, Erik D Roberson²⁰⁵, Gordon Wilcock²⁶⁴, Ekaterina Rogaeva¹²³, Amalia Cecilia Bruni¹¹⁴, Howard J Rosen¹⁶⁴, Maura Gallo¹¹⁴, Roger N Rosenberg²⁶⁶, Yoav Ben-Shlomo²⁶⁷, Mark A Sager¹⁴⁵, Patrizia Mecocci¹²¹, Andrew J Saykin^{139

141}, Pau Pastor^{98

99}, Michael L Cuccaro¹⁴, Jeffery M Vance¹⁴, Julie A Schneider^{153

155

268}, Lori S Schneider^{179

269}, Susan Slifer¹⁴, William W Seeley¹⁶⁴, Amanda G Smith¹⁷³, Joshua A Sonnen⁸⁸, Salvatore Spina⁷⁶, Robert A Stern¹³, Russell H Swerdlow¹⁶⁷, Mitchell Tang¹¹, Rudolph E Tanzi²⁰³, John Q Trojanowski²⁷⁰, Juan C Troncoso²⁷¹, Vivianna M Van Deerlin²⁷⁰, Linda J Van Eldik²⁷², Harry V Vinters^{273

274}, Jean Paul Vonsattel²⁷⁵, Sandra Weintraub^{161

276}, Kathleen A Welsh-Bohmer^{165

277}, Kirk C Wilhelmsen²⁷⁸, Jennifer Williamson²³, Thomas S Wingo^{222

279}, Randall L Woltjer²⁸⁰, Clinton B Wright²⁸¹, Chang-En Yu⁴³, Lei Yu^{153

155}, Yasaman Saba²⁸²; Alzheimer Disease Genetics Consortium (ADGC); European Alzheimer’s Disease Initiative (EADI); Cohorts for Heart and Aging Research in Genomic Epidemiology Consortium (CHARGE); Genetic and Environmental Risk in AD/Defining Genetic, Polygenic and Environmental Risk for Alzheimer’s Disease Consortium (GERAD/PERADES); Alberto Pilotto^{283

284}, Maria J Bullido^{19

285

286}, Oliver Peters^{192

287}, Paul K Crane⁴³, David Bennett^{153

155}, Paola Bosco²⁸⁸, Eliecer Coto²⁵¹, Virginia Boccardi¹²¹, Phil L De Jager²⁸⁹, Alberto Lleo^{19

290}, Nick Warner²⁹¹, Oscar L Lopez^{81

83

158}, Martin Ingelsson⁷⁷, Panagiotis Deloukas²⁹², Carlos Cruchaga^{198

199}, Caroline Graff^{72

73}, Rhian Gwilliam²⁹², Myriam Fornage³⁶, Alison M Goate^{169

293}, Pascual Sanchez-Juan³⁹, Patrick G Kehoe²⁹⁴, Najaf Amin¹⁰, Nilifur Ertekin-Taner^{178

295}, Claudine Berr^{124

125}, Stéphanie Debette^{119

120}, Seth Love²⁹⁴, Lenore J Launer⁶⁶, Steven G Younkin^{178

295}, Jean-Francois Dartigues²⁹⁶, Chris Corcoran²⁹⁷, M Arfan Ikram^{10

298

299}, Dennis W Dickson¹⁷⁸, Gael Nicolas⁴⁴, Dominique Campion^{44

300}, JoAnn Tschanz²⁹⁷, Helena Schmidt^{282

301}, Hakon Hakonarson^{302

303}, Jordi Clarimon^{19

290}, Ron Munger²⁹⁷, Reinhold Schmidt⁴⁹, Lindsay A Farrer^{13

20

212

304

305}, Christine Van Broeckhoven^{15

16}, Michael C O'Donovan⁵, Anita L DeStefano^{13

20}, Lesley Jones^{5

6}, Jonathan L Haines⁵¹, Jean-Francois Deleuze⁹, Michael J Owen⁵, Vilmundur Gudnason^{17

35}, Richard Mayeux^{23

24

25}, Valentina Escott-Price^{5

6}, Bruce M Psaty^{7

32

306

307}, Alfredo Ramirez^{78

191}, Li-San Wang¹¹, Agustin Ruiz^{18

19}, Cornelia M van Duijn¹⁰, Peter A Holmans⁵, Sudha Seshadri^{12

13

308}, Julie Williams^{5

6}, Phillippe Amouyel^{2

3

4

309}, Gerard D Schellenberg^#¹¹, Jean-Charles Lambert^#^{310

311

312}, Margaret A Pericak-Vance^#³¹³

Affiliations

¹ John P. Hussman Institute for Human Genomics, University of Miami Miller School of Medicine, Miami, FL, USA. bkunkle@miami.edu.
² Inserm, U1167, RID-AGE-Risk Factors and Molecular Determinants of Aging-Related Diseases, Lille, France.
³ Institut Pasteur de Lille, Lille, France.
⁴ Univ. Lille, U1167-Excellence Laboratory LabEx DISTALZ, Lille, France.
⁵ Division of Psychological Medicine and Clinical Neurosciences, MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University, Cardiff, UK.
⁶ UK Dementia Research Institute at Cardiff, Cardiff University, Cardiff, UK.
⁷ Cardiovascular Health Research Unit, Department of Medicine, University of Washington, Seattle, WA, USA.
⁸ Department of Biostatistics and Epidemiology/Center for Clinical Epidemiology and Biostatistics, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
⁹ Centre National de Recherche en Génomique Humaine, Institut de Biologie François Jacob, CEA, Université Paris-Saclay, and LabEx GENMED, Evry, France.
¹⁰ Department of Epidemiology, Erasmus Medical Center, Rotterdam, the Netherlands.
¹¹ Penn Neurodegeneration Genomics Center, Department of Pathology and Laboratory Medicine, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
¹² Framingham Heart Study, Framingham, MA, USA.
¹³ Department of Neurology, Boston University School of Medicine, Boston, MA, USA.
¹⁴ John P. Hussman Institute for Human Genomics, University of Miami Miller School of Medicine, Miami, FL, USA.
¹⁵ Neurodegenerative Brain Diseases Group, Center for Molecular Neurology, VIB, Antwerp, Belgium.
¹⁶ Laboratory for Neurogenetics, Institute Born-Bunge, University of Antwerp, Antwerp, Belgium.
¹⁷ Icelandic Heart Association, Kopavogur, Iceland.
¹⁸ Research Center and Memory Clinic of Fundació ACE, Institut Català de Neurociències Aplicades-Universitat Internacional de Catalunya, Barcelona, Spain.
¹⁹ Centro de Investigación Biomédica en Red de Enfermedades Neurodegenerativas, Instituto de Salud Carlos III, Madrid, Spain.
²⁰ Department of Biostatistics, Boston University School of Public Health, Boston, MA, USA.
²¹ Institute of Biomedicine, University of Eastern Finland, Kuopio, Finland.
²² Department of Neurology, Kuopio University Hospital, Kuopio, Finland.
²³ Taub Institute on Alzheimer's Disease and the Aging Brain, Department of Neurology, Columbia University, New York, NY, USA.
²⁴ Gertrude H. Sergievsky Center, Columbia University, New York, NY, USA.
²⁵ Department of Neurology, Columbia University, New York, NY, USA.
²⁶ UMR 894, Center for Psychiatry and Neuroscience, Inserm, Université Paris Descartes, Paris, France.
²⁷ Institute of Human Genetics, University of Bonn, Bonn, Germany.
²⁸ Department of Genomics, Life & Brain Center, University of Bonn, Bonn, Germany.
²⁹ Division of Medical Genetics, University Hospital and Department of Biomedicine, University of Basel, Basel, Switzerland.
³⁰ School of Biotechnology, Dublin City University, Dublin, Ireland.
³¹ Department of Family Medicine, University of Washington, Seattle, WA, USA.
³² Department of Epidemiology, University of Washington, Seattle, WA, USA.
³³ Dementia Research Centre, Department of Neurodegenerative Disease, UCL Institute of Neurology, London, UK.
³⁴ Department of Biostatistics, University of Michigan, Ann Arbor, MI, USA.
³⁵ Faculty of Medicine, University of Iceland, Reykjavik, Iceland.
³⁶ Brown Foundation Institute of Molecular Medicine, University of Texas Health Sciences Center at Houston, Houston, TX, USA.
³⁷ Section of Neuroscience and Clinical Pharmacology, Department of Biomedical Sciences, University of Cagliari, Cagliari, Italy.
³⁸ UK Dementia Research Institute at UCL, Department of Neurodegenerative Disease, UCL Institute of Neurology, London, UK.
³⁹ Neurology Service and CIBERNED, 'Marqués de Valdecilla' University Hospital (University of Cantabria and IDIVAL), Santander, Spain.
⁴⁰ Department of Basic and Clinical Neuroscience, Institute of Psychiatry, Psychology and Neuroscience, King's College London, London, UK.
⁴¹ Department of Immunology, Hospital Universitario Doctor Negrín, Las Palmas de Gran Canaria, Spain.
⁴² Department of Neurology, Medical School, Aristotle University of Thessaloniki, Thessaloniki, Greece.
⁴³ Department of Medicine, University of Washington, Seattle, WA, USA.
⁴⁴ Normandie University, UNIROUEN, Inserm U1245, and Rouen University Hospital, Department of Neurology, Department of Genetics and CNR-MAJ, Normandy Center for Genomic and Personalized Medicine, Rouen, France.
⁴⁵ Department of Neurodegenerative Disease, MRC Prion Unit at UCL, Institute of Prion Diseases, London, UK.
⁴⁶ Centre for Public Health, University of Iceland, Reykjavik, Iceland.
⁴⁷ Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Neurodegenerative Diseases Unit, Milan, Italy.
⁴⁸ University of Milan, Centro Dino Ferrari, Milan, Italy.
⁴⁹ Clinical Division of Neurogeriatrics, Department of Neurology, Medical University Graz, Graz, Austria.
⁵⁰ Institute for Medical Informatics, Statistics and Documentation, Medical University of Graz, Graz, Austria.
⁵¹ Institute for Computational Biology, Department of Population & Quantitative Health Sciences, Case Western Reserve University, Cleveland, OH, USA.
⁵² Department of Psychiatry and Psychotherapy, University of Erlangen-Nuremberg, Erlangen, Germany.
⁵³ Program in Medical and Population Genetics, Broad Institute, Cambridge, MA, USA.
⁵⁴ Department of Pharmacology and Neuroscience, University of North Texas Health Science Center, Fort Worth, TX, USA.
⁵⁵ Laboratory for Neurochemistry and Behavior, Institute Born-Bunge, University of Antwerp, Antwerp, Belgium.
⁵⁶ Department of Neurology and Memory Clinic, Hospital Network Antwerp, Antwerp, Belgium.
⁵⁷ Department of Psychiatry and Psychotherapy, University Hospital, Saarland, Germany.
⁵⁸ Department of Psychiatry, University of Texas Southwestern Medical Center, Dallas, TX, USA.
⁵⁹ Laboratory for Cognitive Neurology, Department of Neurology, University Hospital and University of Leuven, Leuven, Belgium.
⁶⁰ Department of Neurology, Johns Hopkins University, Baltimore, MD, USA.
⁶¹ National Alzheimer's Coordinating Center, University of Washington, Seattle, WA, USA.
⁶² Department of Psychiatry, Martin Luther University Halle-Wittenberg, Halle, Germany.
⁶³ Department of Epidemiology, Harvard T.H. Chan School of Public Health, Harvard University, Boston, MA, USA.
⁶⁴ Department of Psychiatry, Massachusetts General Hospital/Harvard Medical School, Boston, MA, USA.
⁶⁵ Department of Psychiatry, University of Oxford, Oxford, UK.
⁶⁶ Laboratory of Epidemiology and Population Sciences, National Institute on Aging, Bethesda, MD, USA.
⁶⁷ Alzheimer's Disease and Memory Disorders Center, Baylor College of Medicine, Houston, TX, USA.
⁶⁸ Division of Clinical Neurosciences, School of Medicine, University of Southampton, Southampton, UK.
⁶⁹ Section of Computational Biomedicine, Department of Medicine, Boston University School of Medicine, Boston, MA, USA.
⁷⁰ Office of Strategy and Measurement, University of North Texas Health Science Center, Fort Worth, TX, USA.
⁷¹ C.S. Kubik Laboratory for Neuropathology, Massachusetts General Hospital, Charlestown, MA, USA.
⁷² Theme Aging, Unit for Hereditary Dementias, Karolinska University Hospital-Solna, Stockholm, Sweden.
⁷³ Karolinska Institutet, Department of Neurobiology, Care Sciences and Society, Alzheimer Research Center, Division of Neurogeriatrics, Solna, Sweden.
⁷⁴ German Center for Neurodegenerative Diseases, Bonn, Germany.
⁷⁵ Department of Psychiatry and Psychotherapy, University of Bonn, Bonn, Germany.
⁷⁶ Department of Pathology and Laboratory Medicine, Indiana University, Indianapolis, IN, USA.
⁷⁷ Department of Public Health and Caring Sciences/Geriatrics, Uppsala University, Uppsala, Sweden.
⁷⁸ Department for Neurodegenerative Diseases and Geriatric Psychiatry, University Hospital Bonn, Bonn, Germany.
⁷⁹ Department of Surgery, University of Texas Southwestern Medical Center, Dallas, TX, USA.
⁸⁰ Institute for Medical Informatics, Biometry and Epidemiology, University Hospital of Essen, University Duisburg-Essen, Essen, Germany.
⁸¹ Department of Psychiatry, University of Pittsburgh, Pittsburgh, PA, USA.
⁸² Department of Human Genetics, University of Pittsburgh, Pittsburgh, PA, USA.
⁸³ Alzheimer's Disease Research Center, University of Pittsburgh, Pittsburgh, PA, USA.
⁸⁴ Institute of Genetics, Queen's Medical Centre, University of Nottingham, Nottingham, UK.
⁸⁵ Department of Neurology, Albert Einstein College of Medicine, Bronx, NY, USA.
⁸⁶ Section of Neuroscience, DIMEC-University of Parma, Parma, Italy.
⁸⁷ FERB-Alzheimer Center, Gazzaniga (Bergamo), Italy.
⁸⁸ Department of Pathology, University of Washington, Seattle, WA, USA.
⁸⁹ Elderly and Psychiatric Disorders Department, Medical University of Lodz, Lodz, Poland.
⁹⁰ Mercer's Institute for Research on Aging, St. James's Hospital and Trinity College, Dublin, Ireland.
⁹¹ St. James's Hospital and Trinity College, Dublin, Ireland.
⁹² Kaiser Permanente Washington Health Research Institute, Seattle, WA, USA.
⁹³ A.I. Virtanen Institute for Molecular Sciences, University of Eastern Finland, Kuopio, Finland.
⁹⁴ Departments of Medicine, Geriatrics, Gerontology and Neurology, University of Mississippi Medical Center, Jackson, MS, USA.
⁹⁵ Interdisciplinary Department of Medicine, Geriatric Medicine and Memory Unity, University of Bari, Bari, Italy.
⁹⁶ Department of Neurology, University of Washington, Seattle, WA, USA.
⁹⁷ Department of Geriatrics, Center for Aging Brain, University of Bari, Bari, Italy.
⁹⁸ Fundació per la Recerca Biomèdica i Social Mútua Terrassa, Terrassa, Barcelona, Spain.
⁹⁹ Memory Disorders Unit, Department of Neurology, Hospital Universitari Mutua de Terrassa, Terrassa, Barcelona, Spain.
¹⁰⁰ Department of Internal Medicine, Erasmus University Medical Center, Rotterdamt, the Netherlands.
¹⁰¹ Netherlands Consortium on Health Aging and National Genomics Initiative, Leiden, the Netherlands.
¹⁰² Department of Neurology, Mayo Clinic, Rochester, MN, USA.
¹⁰³ CHU Lille, Memory Center of Lille (Centre Mémoire de Ressources et de Recherche), Lille, France.
¹⁰⁴ Department of Clinical and Behavioral Neurology, Experimental Neuropsychobiology Laboratory, IRCCS Santa Lucia Foundation, Rome, Italy.
¹⁰⁵ School of Public Health, Human Genetics Center, University of Texas Health Science Center at Houston, Houston, TX, USA.
¹⁰⁶ Human Genome Sequencing Center, Baylor College of Medicine, Houston, TX, USA.
¹⁰⁷ Neurogenomics Division, Translational Genomics Research Institute, Phoenix, AZ, USA.
¹⁰⁸ Arizona Alzheimer's Consortium, Phoenix, AZ, USA.
¹⁰⁹ Banner Alzheimer's Institute, Phoenix, AZ, USA.
¹¹⁰ Department of Psychiatry, University of Arizona, Phoenix, AZ, USA.
¹¹¹ Institute for Translational Genomics and Population Sciences, Departments of Pediatrics and Medicine, Los Angeles BioMedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.
¹¹² Department of Clinical Sciences, University of Texas Southwestern Medical Center, Dallas, TX, USA.
¹¹³ University Paris Descartes, EA 4468, AP-HP, Geriatrics Department, Hôpital Broca, Paris, France.
¹¹⁴ Regional Neurogenetic Centre (CRN), ASP Catanzaro, Lamezia Terme, Italy.
¹¹⁵ Departments of Psychiatry, Medicine, Family & Community Medicine, South Texas Veterans Health Administration Geriatric Research Education & Clinical Center (GRECC), UT Health Science Center at San Antonio, San Antonio, TX, USA.
¹¹⁶ University of Bordeaux, Inserm 1219, Bordeaux, France.
¹¹⁷ Department of Neurology, Bordeaux University Hospital / CHU de Bordeaux, Bordeaux, France.
¹¹⁸ Department of Psychiatry, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
¹¹⁹ Inserm U1127, CNRS UMR 7225, Sorbonne Universités, UPMC Université Paris 06, UMRS 1127, Institut du Cerveau et de la Moelle Épinière, Paris, France.
¹²⁰ AP-HP, Department of Genetics, Pitié-Salpêtrière Hospital, Paris, France.
¹²¹ Section of Gerontology and Geriatrics, Department of Medicine, University of Perugia, Perugia, Italy.
¹²² Cambridge Institute for Medical Research, University of Cambridge, Cambridge, UK.
¹²³ Tanz Centre for Research in Neurodegenerative Disease, University of Toronto, Toronto, Ontario, Canada.
¹²⁴ Inserm U1061 Neuropsychiatry, La Colombière Hospital, Montpellier, France.
¹²⁵ Montpellier University, Montpellier, France.
¹²⁶ Department of Clinical Brain Sciences, University of Edinburgh, Edinburgh, UK.
¹²⁷ VA Puget Sound Health Care System/>GRECC, Seattle, WA, USA.
¹²⁸ Department of Psychiatry and Behavioral Sciences, University of Washington School of Medicine, Seattle, WA, USA.
¹²⁹ Institut de la Mémoire et de la Maladie d'Alzheimer and Institut du Cerveau et de la Moelle Épinière, Département de Neurologie, Hôpital de la Pitié-Salpêtrière, Paris, France.
¹³⁰ Institut des Neurosciences Translationnelles de Paris, Institut du Cerveau et de la Moelle Épinière, Paris, France.
¹³¹ Inserm, CNRS, UMR-S975, Institut du Cerveau et de la Moelle Epinière, Paris, France.
¹³² Sorbonne Universités, Université Pierre et Marie Curie, Hôpital de la Pitié-Salpêtrière, AP-HP, Paris, France.
¹³³ Ageing Group, Centre for Public Health, School of Medicine, Dentistry and Biomedical Sciences, Queen's University Belfast, Belfast, UK.
¹³⁴ Departments of Neurology, Pharmacology & Neuroscience, Texas Tech University Health Science Center, Lubbock, TX, USA.
¹³⁵ Department of Neurology, University of Michigan, Ann Arbor, MI, USA.
¹³⁶ Geriatric Research, Education and Clinical Center (GRECC), VA Ann Arbor Healthcare System (VAAAHS), Ann Arbor, MI, USA.
¹³⁷ Michigan Alzheimer Disease Center, Ann Arbor, MI, USA.
¹³⁸ Aging Research Center, Department of Neurobiology, Care Sciences and Society, Karolinska Institutet and Stockholm University, Stockholm, Sweden.
¹³⁹ Indiana Alzheimer's Disease Center, Indiana University School of Medicine, Indianapolis, IN, USA.
¹⁴⁰ Department of Medical and Molecular Genetics, Indiana University, Indianapolis, IN, USA.
¹⁴¹ Department of Neurology, Indiana University, Indianapolis, IN, USA.
¹⁴² Department of Radiology and Imaging Sciences, Indiana University, Indianapolis, IN, USA.
¹⁴³ Department of Psychiatry, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
¹⁴⁴ Geriatric Research, Education and Clinical Center (GRECC), University of Wisconsin, Madison, WI, USA.
¹⁴⁵ Department of Medicine, University of Wisconsin, Madison, WI, USA.
¹⁴⁶ Wisconsin Alzheimer's Disease Research Center, Madison, WI, USA.
¹⁴⁷ AXA Research Fund & Sorbonne University Chair, Paris, France.
¹⁴⁸ Sorbonne University, GRC n° 21, Alzheimer Precision Medicine (APM), AP-HP, Pitié-Salpêtrière Hospital, Paris, France.
¹⁴⁹ Brain & Spine Institute, Inserm U 1127, CNRS UMR 7225, Paris, France.
¹⁵⁰ Institute of Memory and Alzheimer's Disease, Department of Neurology, Pitié-Salpêtrière Hospital, AP-HP, Paris, France.
¹⁵¹ Grupo de Medicina Xenomica, Universidade de Santiago de Compostela, Centro Nacional de Genotipado, Centro de Investigación Biomédica en Red de Enfermedades Raras, Santiago de Compostela, Spain.
¹⁵² UK Dementia Research Institute, University of Cambridge, Cambridge, UK.
¹⁵³ Department of Neurological Sciences, Rush University Medical Center, Chicago, IL, USA.
¹⁵⁴ Department of Behavioral Sciences, Rush University Medical Center, Chicago, IL, USA.
¹⁵⁵ Rush Alzheimer's Disease Center, Rush University Medical Center, Chicago, IL, USA.
¹⁵⁶ Central Institute of Mental Health, Medical Faculty Mannheim, University of Heidelberg, Heidelberg, Germany.
¹⁵⁷ Civin Laboratory for Neuropathology, Banner Sun Health Research Institute, Phoenix, AZ, USA.
¹⁵⁸ Department of Neurology, University of Pittsburgh, Pittsburgh, PA, USA.
¹⁵⁹ Department of Psychology, University of Pittsburgh School of Medicine, Pittsburgh, PA, USA.
¹⁶⁰ Department of Pathology, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
¹⁶¹ Mesulam Center for Cognitive Neurology and Alzheimer's Disease, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
¹⁶² Genetic Epidemiology, QIMR Berghofer Medical Research Institute, Herston, Queensland, Australia.
¹⁶³ Swedish Medical Center, Seattle, WA, USA.
¹⁶⁴ Department of Neurology, University of California, San Francisco, San Francisco, CA, USA.
¹⁶⁵ Department of Neurology, Duke University, Durham, NC, USA.
¹⁶⁶ Departments of Biology, Brigham Young University, Provo, UT, USA.
¹⁶⁷ University of Kansas Alzheimer's Disease Center, University of Kansas Medical Center, Kansas City, KS, USA.
¹⁶⁸ Department of Experimental and Clinical Medicine, Neurological Institute, University of Pisa, Pisa, Italy.
¹⁶⁹ Department of Genetics and Genomic Sciences, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
¹⁷⁰ Department of Neuroscience, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
¹⁷¹ Department of Pathology and Immunology, Washington University, St. Louis, MO, USA.
¹⁷² Division of Psychiatry, University College London, London, UK.
¹⁷³ USF Health Byrd Alzheimer's Institute, University of South Florida, Tampa, FL, USA.
¹⁷⁴ Fred Hutchinson Cancer Research Center, Seattle, WA, USA.
¹⁷⁵ Department of Molecular Neuroscience, UCL, Institute of Neurology, London, UK.
¹⁷⁶ Mental Health & Behavioral Science Service, Bruce W. Carter VA Medical Center, Miami, FL, USA.
¹⁷⁷ Department of Neurodegenerative Disease, UCL Institute of Neurology, London, UK.
¹⁷⁸ Department of Neuroscience, Mayo Clinic, Jacksonville, FL, USA.
¹⁷⁹ Department of Neurology, University of Southern California, Los Angeles, CA, USA.
¹⁸⁰ Department of Neurology, Catholic University of Rome, Rome, Italy.
¹⁸¹ Department of Psychiatry and Behavioral Sciences, Miller School of Medicine, University of Miami, Miami, FL, USA.
¹⁸² Department of Neurology, University of California, Davis, Sacramento, CA, USA.
¹⁸³ Institute for Memory Impairments and Neurological Disorders, University of California, Irvine, Irvine, CA, USA.
¹⁸⁴ Wien Center for Alzheimer's Disease and Memory Disorders, Mount Sinai Medical Center, Miami Beach, FL, USA.
¹⁸⁵ Rush Institute for Healthy Aging, Department of Internal Medicine, Rush University Medical Center, Chicago, IL, USA.
¹⁸⁶ Department of Old Age Psychiatry, Institute of Psychiatry, Psychology and Neuroscience, King's College London, London, UK.
¹⁸⁷ Department of Primary Medical Care, University Medical Centre Hamburg-Eppendorf, Hamburg, Germany.
¹⁸⁸ Department of Pathology, University of Alabama at Birmingham, Birmingham, AL, USA.
¹⁸⁹ Sanders-Brown Center on Aging, Department of Biostatistics, University of Kentucky, Lexington, KY, USA.
¹⁹⁰ Department of Psychiatry, New York University, New York, NY, USA.
¹⁹¹ Department of Psychiatry and Psychotherapy, University of Cologne, Cologne, Germany.
¹⁹² Department of Psychiatry and Psychotherapy, Charité University Medicine, Berlin, Germany.
¹⁹³ Department of Neurosciences, University of California, San Diego, La Jolla, CA, USA.
¹⁹⁴ Department of Pathology and Laboratory Medicine, Emory University, Atlanta, GA, USA.
¹⁹⁵ Emory Alzheimer's Disease Center, Emory University, Atlanta, GA, USA.
¹⁹⁶ Department of Psychiatry, University of Freiburg, Freiburg, Germany.
¹⁹⁷ Neurogenetics Program, University of California, Los Angeles, Los Angeles, CA, USA.
¹⁹⁸ Department of Psychiatry, Washington University School of Medicine, St. Louis, MO, USA.
¹⁹⁹ Hope Center Program on Protein Aggregation and Neurodegeneration, Washington University School of Medicine, St. Louis, MO, USA.
²⁰⁰ Division of Genetics, Department of Medicine and Partners Center for Personalized Genetic Medicine, Brigham and Women's Hospital and Harvard Medical School, Boston, MA, USA.
²⁰¹ Department of Neurology, Washington University, St. Louis, MO, USA.
²⁰² Department of Genetics, Washington University, St. Louis, MO, USA.
²⁰³ Department of Neurology, Massachusetts General Hospital/Harvard Medical School, Boston, MA, USA.
²⁰⁴ Department of Pathology (Neuropathology), University of Pittsburgh, Pittsburgh, PA, USA.
²⁰⁵ Department of Neurology, University of Alabama at Birmingham, Birmingham, AL, USA.
²⁰⁶ Cologne Center for Genomics, University of Cologne, Cologne, Germany.
²⁰⁷ Department of Pathology, Duke University, Durham, NC, USA.
²⁰⁸ Department of Genome Sciences, University of Washington, Seattle, WA, USA.
²⁰⁹ Department of Medicine (Medical Genetics), University of Washington, Seattle, WA, USA.
²¹⁰ Sanders-Brown Center on Aging, College of Public Health, Department of Epidemiology, University of Kentucky, Lexington, KY, USA.
²¹¹ Unidad Clínica de Enfermedades Infecciosas y Microbiología, Hospital Universitario de Valme, Sevilla, Spain.
²¹² Department of Medicine (Biomedical Genetics), Boston University School of Medicine, Boston, MA, USA.
²¹³ Functional Genomics Center Zurich, ETH/University of Zurich, Zurich, Switzerland.
²¹⁴ Department of Neurology, Oregon Health &Science University, Portland, OR, USA.
²¹⁵ Department of Neurology, Portland Veterans Affairs Medical Center, Portland, OR, USA.
²¹⁶ Department of Pathology and Laboratory Medicine, University of California, Irvine, Irvine, CA, USA.
²¹⁷ Department of Pathology, Boston University School of Medicine, Boston University, Boston, MA, USA.
²¹⁸ Inserm U558, University of Toulouse, Toulouse, France.
²¹⁹ Department of Neuropsychology, University of California San Francisco, San Francisco, CA, USA.
²²⁰ Institute for Ageing and Health, Newcastle University, Newcastle upon Tyne, UK.
²²¹ Department of Neurobiology and Behavior, University of California, Irvine, Irvine, CA, USA.
²²² Department of Neurology, Emory University, Atlanta, GA, USA.
²²³ Institute for Stroke and Dementia Research, Klinikum der Universität München, Munich, Germany.
²²⁴ German Center for Neurodegenerative Diseases, Munich, Germany.
²²⁵ Cleveland Clinic Lou Ruvo Center for Brain Health, Cleveland Clinic, Cleveland, OH, USA.
²²⁶ Division of Neuroscience and Experimental Psychology, School of Biological Sciences, Faculty of Biology, Medicine and Health, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK.
²²⁷ Department of Pathology, University of Michigan, Ann Arbor, MI, USA.
²²⁸ Institute of Epidemiology, Helmholtz Zentrum München, German Research Center for Environmental Health, Neuherberg, Munich, Germany.
²²⁹ Helmholtz Center Munich, Institute of Epidemiology, Neuherberg, Munich, Germany.
²³⁰ Ludwig-Maximilians University Chair of Epidemiology, Munich, Germany.
²³¹ Joint Biobank Munich and KORA Biobank, Baltimore, MD, USA.
²³² Department of Psychiatry, Johns Hopkins University, Baltimore, MD, USA.
²³³ Human Genetics, Schools of Life Sciences and Medicine, University of Nottingham, Nottingham, UK.
²³⁴ Department of Medicine-Pulmonary, New York University, New York, NY, USA.
²³⁵ Department of Neurology, University of Miami, Miami, FL, USA.
²³⁶ Department of Pathology, University of California, San Diego, La Jolla, CA, USA.
²³⁷ Institute of Neurology, Catholic University of Sacred Hearth, Rome, Italy.
²³⁸ School of Nursing Northwest Research Group on Aging, University of Washington, Seattle, WA, USA.
²³⁹ Institute of Primary Care and Public Health, Cardiff University, University Hospital of Wales, Cardiff, UK.
²⁴⁰ Department of Neurology, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
²⁴¹ Cambridge Institute of Public Health, University of Cambridge School of Clinical Medicine, Cambridge, UK.
²⁴² Weill Institute for Neurosciences, Memory and Aging Center, University of California, San Francisco, San Francisco, CA, USA.
²⁴³ Institute of Social Medicine, Occupational Health and Public Health, University of Leipzig, Leipzig, Germany.
²⁴⁴ Department of Pathology, University of Southern California, Los Angeles, CA, USA.
²⁴⁵ Department of Pathology and Laboratory Medicine, University of California, Davis, Sacramento, CA, USA.
²⁴⁶ Institute of Psychiatry, Psychology and Neuroscienceó, King's College London, London, UK.
²⁴⁷ UK Dementia Research Institute, King's College London, London, UK.
²⁴⁸ Department of Psychiatry and Psychotherapy, University Medical Center Goettingen, Goettingen, Germany.
²⁴⁹ German Center for Neurodegenerative Diseases, Goettingen, Germany.
²⁵⁰ IBiMED, Medical Sciences Department, University of Aveiro, Aveiro, Portugal.
²⁵¹ Molecular Genetics Laboratory-Hospital, University of Central Asturias, Oviedo, Spain.
²⁵² Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, MN, USA.
²⁵³ Molecular Genetics Section, Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, MD, USA.
²⁵⁴ Department of Neurology, University of California, Irvine, Irvine, CA, USA.
²⁵⁵ Department of Neuroscience, Psychology, Drug Research and Child Health, University of Florence, Florence, Italy.
²⁵⁶ Centro di Ricerca, Trasferimento e Alta Formazione DENOTHE, University of Florence, Florence, Italy.
²⁵⁷ Department of Neurology, University of Colorado School of Medicine, Aurora, CO, USA.
²⁵⁸ IRCCS Fondazione Don Carlo Gnocchi, Florence, Italy.
²⁵⁹ Laboratory of Neuropsychiatry, IRCCS Santa Lucia Foundation, Rome, Italy.
²⁶⁰ Division of Neuropsychiatry, Department of Psychiatry and Behavioral Sciences, Baylor College of Medicine, Houston, TX, USA.
²⁶¹ Alzheimer's Disease Center, New York University, New York, NY, USA.
²⁶² Oxford Healthy Aging Project, Clinical Trial Service Unit, University of Oxford, Oxford, UK.
²⁶³ Department of Epidemiology, Columbia University, New York, NY, USA.
²⁶⁴ Oxford Project to Investigate Memory and Ageing, University of Oxford, Nuffield Department of Clinical Neurosciences, John Radcliffe Hospital, Oxford, UK.
²⁶⁵ Department of Neurology, Keck School of Medicine at the University of Southern California, Los Angeles, Los Angeles, CA, USA.
²⁶⁶ Department of Neurology, University of Texas Southwestern Medical Center, Dallas, TX, USA.
²⁶⁷ Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK.
²⁶⁸ Department of Pathology (Neuropathology), Rush University Medical Center, Chicago, IL, USA.
²⁶⁹ Department of Psychiatry, University of Southern California, Los Angeles, CA, USA.
²⁷⁰ Department of Pathology and Laboratory Medicine, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
²⁷¹ Department of Pathology, Johns Hopkins University, Baltimore, MD, USA.
²⁷² Sanders-Brown Center on Aging, Department of Neuroscience, University of Kentucky, Lexington, KY, USA.
²⁷³ Department of Neurology, University of California, Los Angeles, Los Angeles, CA, USA.
²⁷⁴ Department of Pathology and Laboratory Medicine, University of California, Los Angeles, Los Angeles, CA, USA.
²⁷⁵ Taub Institute on Alzheimer's Disease and the Aging Brain, Department of Pathology, Columbia University, New York, NY, USA.
²⁷⁶ Department of Psychiatry and Behavioral Sciences, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
²⁷⁷ Department of Psychiatry and Behavioral Sciences, Duke University, Durham, NC, USA.
²⁷⁸ Department of Genetics, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.
²⁷⁹ Department of Human Genetics, Emory University, Atlanta, GA, USA.
²⁸⁰ Department of Pathology, Oregon Health & Science University, Portland, OR, USA.
²⁸¹ National Institute of Neurological Disorders and Stroke, Bethesda, MD, USA.
²⁸² Gottfried Schatz Research Center for Cell Signaling, Metabolism and Aging, Division of Molecular Biology and Biochemistry, Medical University Graz, Graz, Austria.
²⁸³ Gerontology and Geriatrics Research Laboratory, IRCCS Casa Sollievo della Sofferenza, San Giovanni Rotondo, Italy.
²⁸⁴ Department Geriatric Care, Orthogeriatrics and Rehabilitation, Galliera Hospital, Genova, Italy.
²⁸⁵ IdiPAZ, Instituto de Investigación Sanitaria la Paz, Madrid, Spain.
²⁸⁶ Centro de Biologia Molecular Severo Ochoa (CSIC-UAM), Madrid, Spain.
²⁸⁷ German Center for Neurodegenerative Diseases, Berlin, Germany.
²⁸⁸ Instituto di Ricovero e Cura a Carattere Scientifico, Associazione Oasi Maria Santissima Srl, Troina, Italy.
²⁸⁹ Center for Translational and Computational Neuroimmunology, Department of Neurology, Columbia University Medical Center, New York, NY, USA.
²⁹⁰ Memory Unit, Neurology Department and Sant Pau Biomedical Research Institute, Hospital de la Santa Creu i Sant Pau, Autonomous University Barcelona, Barcelona, Spain.
²⁹¹ Somerset Partnership NHS Trust, Somerset, UK.
²⁹² The Wellcome Trust Sanger Institute, Hinxton, Cambridge, UK.
²⁹³ Ronald M. Loeb Center for Alzheimer's Disease, Department of Neuroscience, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
²⁹⁴ University of Bristol Medical School, Learning & Research level 2, Southmead Hospital, Bristol, UK.
²⁹⁵ Department of Neurology, Mayo Clinic, Jacksonville, FL, USA.
²⁹⁶ Memory Research and Resources Center, CMRR de Bordeaux, Bordeaux, France.
²⁹⁷ Utah State University, Logan, UT, USA.
²⁹⁸ Department of Neurology, Erasmus MC University Medical Center, Rotterdam, the Netherlands.
²⁹⁹ Departments of Radiology, Erasmus MC University Medical Center, Rotterdam, the Netherlands.
³⁰⁰ Department of Research Rouvray Psychiatric Hospital, Sotteville-lès-Rouen, France.
³⁰¹ Department of Neurology, Medical University Graz, Graz, Austria.
³⁰² Center for Applied Genomics, Children's Hospital of Philadelphia, The Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA.
³⁰³ Division of Human Genetics, Department of Pediatrics, The Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA.
³⁰⁴ Department of Ophthalmology, Boston University School of Medicine, Boston University, Boston, MA, USA.
³⁰⁵ Department of Epidemiology, Boston University School of Public Health, Boston, MA, USA.
³⁰⁶ Department of Health Services, University of Washington, Seattle, WA, USA.
³⁰⁷ Kaiser Permanente, Washington Health Research Institute, Seattle, WA, USA.
³⁰⁸ Glenn Biggs Institute for Alzheimer's and Neurodegenerative Diseases, San Antonio, TX, USA.
³⁰⁹ Centre Hospitalier Universitaire de Lille, Lille, France.
³¹⁰ Inserm, U1167, RID-AGE-Risk Factors and Molecular Determinants of Aging-Related Diseases, Lille, France. jean-charles.lambert@pasteur-lille.fr.
³¹¹ Institut Pasteur de Lille, Lille, France. jean-charles.lambert@pasteur-lille.fr.
³¹² Univ. Lille, U1167-Excellence Laboratory LabEx DISTALZ, Lille, France. jean-charles.lambert@pasteur-lille.fr.
³¹³ John P. Hussman Institute for Human Genomics, University of Miami Miller School of Medicine, Miami, FL, USA. mpericak@miami.edu.

^# Contributed equally.

PMID: 31417202
PMCID: PMC7265117
DOI: 10.1038/s41588-019-0495-7

Published Erratum

Author Correction: Genetic meta-analysis of diagnosed Alzheimer's disease identifies new risk loci and implicates Aβ, tau, immunity and lipid processing

Brian W Kunkle et al. Nat Genet. 2019 Sep.

. 2019 Sep;51(9):1423-1424.

doi: 10.1038/s41588-019-0495-7.

Authors

Affiliations

¹ John P. Hussman Institute for Human Genomics, University of Miami Miller School of Medicine, Miami, FL, USA. bkunkle@miami.edu.
² Inserm, U1167, RID-AGE-Risk Factors and Molecular Determinants of Aging-Related Diseases, Lille, France.
³ Institut Pasteur de Lille, Lille, France.
⁴ Univ. Lille, U1167-Excellence Laboratory LabEx DISTALZ, Lille, France.
⁵ Division of Psychological Medicine and Clinical Neurosciences, MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University, Cardiff, UK.
⁶ UK Dementia Research Institute at Cardiff, Cardiff University, Cardiff, UK.
⁷ Cardiovascular Health Research Unit, Department of Medicine, University of Washington, Seattle, WA, USA.
⁸ Department of Biostatistics and Epidemiology/Center for Clinical Epidemiology and Biostatistics, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
⁹ Centre National de Recherche en Génomique Humaine, Institut de Biologie François Jacob, CEA, Université Paris-Saclay, and LabEx GENMED, Evry, France.
¹⁰ Department of Epidemiology, Erasmus Medical Center, Rotterdam, the Netherlands.
¹¹ Penn Neurodegeneration Genomics Center, Department of Pathology and Laboratory Medicine, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
¹² Framingham Heart Study, Framingham, MA, USA.
¹³ Department of Neurology, Boston University School of Medicine, Boston, MA, USA.
¹⁴ John P. Hussman Institute for Human Genomics, University of Miami Miller School of Medicine, Miami, FL, USA.
¹⁵ Neurodegenerative Brain Diseases Group, Center for Molecular Neurology, VIB, Antwerp, Belgium.
¹⁶ Laboratory for Neurogenetics, Institute Born-Bunge, University of Antwerp, Antwerp, Belgium.
¹⁷ Icelandic Heart Association, Kopavogur, Iceland.
¹⁸ Research Center and Memory Clinic of Fundació ACE, Institut Català de Neurociències Aplicades-Universitat Internacional de Catalunya, Barcelona, Spain.
¹⁹ Centro de Investigación Biomédica en Red de Enfermedades Neurodegenerativas, Instituto de Salud Carlos III, Madrid, Spain.
²⁰ Department of Biostatistics, Boston University School of Public Health, Boston, MA, USA.
²¹ Institute of Biomedicine, University of Eastern Finland, Kuopio, Finland.
²² Department of Neurology, Kuopio University Hospital, Kuopio, Finland.
²³ Taub Institute on Alzheimer's Disease and the Aging Brain, Department of Neurology, Columbia University, New York, NY, USA.
²⁴ Gertrude H. Sergievsky Center, Columbia University, New York, NY, USA.
²⁵ Department of Neurology, Columbia University, New York, NY, USA.
²⁶ UMR 894, Center for Psychiatry and Neuroscience, Inserm, Université Paris Descartes, Paris, France.
²⁷ Institute of Human Genetics, University of Bonn, Bonn, Germany.
²⁸ Department of Genomics, Life & Brain Center, University of Bonn, Bonn, Germany.
²⁹ Division of Medical Genetics, University Hospital and Department of Biomedicine, University of Basel, Basel, Switzerland.
³⁰ School of Biotechnology, Dublin City University, Dublin, Ireland.
³¹ Department of Family Medicine, University of Washington, Seattle, WA, USA.
³² Department of Epidemiology, University of Washington, Seattle, WA, USA.
³³ Dementia Research Centre, Department of Neurodegenerative Disease, UCL Institute of Neurology, London, UK.
³⁴ Department of Biostatistics, University of Michigan, Ann Arbor, MI, USA.
³⁵ Faculty of Medicine, University of Iceland, Reykjavik, Iceland.
³⁶ Brown Foundation Institute of Molecular Medicine, University of Texas Health Sciences Center at Houston, Houston, TX, USA.
³⁷ Section of Neuroscience and Clinical Pharmacology, Department of Biomedical Sciences, University of Cagliari, Cagliari, Italy.
³⁸ UK Dementia Research Institute at UCL, Department of Neurodegenerative Disease, UCL Institute of Neurology, London, UK.
³⁹ Neurology Service and CIBERNED, 'Marqués de Valdecilla' University Hospital (University of Cantabria and IDIVAL), Santander, Spain.
⁴⁰ Department of Basic and Clinical Neuroscience, Institute of Psychiatry, Psychology and Neuroscience, King's College London, London, UK.
⁴¹ Department of Immunology, Hospital Universitario Doctor Negrín, Las Palmas de Gran Canaria, Spain.
⁴² Department of Neurology, Medical School, Aristotle University of Thessaloniki, Thessaloniki, Greece.
⁴³ Department of Medicine, University of Washington, Seattle, WA, USA.
⁴⁴ Normandie University, UNIROUEN, Inserm U1245, and Rouen University Hospital, Department of Neurology, Department of Genetics and CNR-MAJ, Normandy Center for Genomic and Personalized Medicine, Rouen, France.
⁴⁵ Department of Neurodegenerative Disease, MRC Prion Unit at UCL, Institute of Prion Diseases, London, UK.
⁴⁶ Centre for Public Health, University of Iceland, Reykjavik, Iceland.
⁴⁷ Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Neurodegenerative Diseases Unit, Milan, Italy.
⁴⁸ University of Milan, Centro Dino Ferrari, Milan, Italy.
⁴⁹ Clinical Division of Neurogeriatrics, Department of Neurology, Medical University Graz, Graz, Austria.
⁵⁰ Institute for Medical Informatics, Statistics and Documentation, Medical University of Graz, Graz, Austria.
⁵¹ Institute for Computational Biology, Department of Population & Quantitative Health Sciences, Case Western Reserve University, Cleveland, OH, USA.
⁵² Department of Psychiatry and Psychotherapy, University of Erlangen-Nuremberg, Erlangen, Germany.
⁵³ Program in Medical and Population Genetics, Broad Institute, Cambridge, MA, USA.
⁵⁴ Department of Pharmacology and Neuroscience, University of North Texas Health Science Center, Fort Worth, TX, USA.
⁵⁵ Laboratory for Neurochemistry and Behavior, Institute Born-Bunge, University of Antwerp, Antwerp, Belgium.
⁵⁶ Department of Neurology and Memory Clinic, Hospital Network Antwerp, Antwerp, Belgium.
⁵⁷ Department of Psychiatry and Psychotherapy, University Hospital, Saarland, Germany.
⁵⁸ Department of Psychiatry, University of Texas Southwestern Medical Center, Dallas, TX, USA.
⁵⁹ Laboratory for Cognitive Neurology, Department of Neurology, University Hospital and University of Leuven, Leuven, Belgium.
⁶⁰ Department of Neurology, Johns Hopkins University, Baltimore, MD, USA.
⁶¹ National Alzheimer's Coordinating Center, University of Washington, Seattle, WA, USA.
⁶² Department of Psychiatry, Martin Luther University Halle-Wittenberg, Halle, Germany.
⁶³ Department of Epidemiology, Harvard T.H. Chan School of Public Health, Harvard University, Boston, MA, USA.
⁶⁴ Department of Psychiatry, Massachusetts General Hospital/Harvard Medical School, Boston, MA, USA.
⁶⁵ Department of Psychiatry, University of Oxford, Oxford, UK.
⁶⁶ Laboratory of Epidemiology and Population Sciences, National Institute on Aging, Bethesda, MD, USA.
⁶⁷ Alzheimer's Disease and Memory Disorders Center, Baylor College of Medicine, Houston, TX, USA.
⁶⁸ Division of Clinical Neurosciences, School of Medicine, University of Southampton, Southampton, UK.
⁶⁹ Section of Computational Biomedicine, Department of Medicine, Boston University School of Medicine, Boston, MA, USA.
⁷⁰ Office of Strategy and Measurement, University of North Texas Health Science Center, Fort Worth, TX, USA.
⁷¹ C.S. Kubik Laboratory for Neuropathology, Massachusetts General Hospital, Charlestown, MA, USA.
⁷² Theme Aging, Unit for Hereditary Dementias, Karolinska University Hospital-Solna, Stockholm, Sweden.
⁷³ Karolinska Institutet, Department of Neurobiology, Care Sciences and Society, Alzheimer Research Center, Division of Neurogeriatrics, Solna, Sweden.
⁷⁴ German Center for Neurodegenerative Diseases, Bonn, Germany.
⁷⁵ Department of Psychiatry and Psychotherapy, University of Bonn, Bonn, Germany.
⁷⁶ Department of Pathology and Laboratory Medicine, Indiana University, Indianapolis, IN, USA.
⁷⁷ Department of Public Health and Caring Sciences/Geriatrics, Uppsala University, Uppsala, Sweden.
⁷⁸ Department for Neurodegenerative Diseases and Geriatric Psychiatry, University Hospital Bonn, Bonn, Germany.
⁷⁹ Department of Surgery, University of Texas Southwestern Medical Center, Dallas, TX, USA.
⁸⁰ Institute for Medical Informatics, Biometry and Epidemiology, University Hospital of Essen, University Duisburg-Essen, Essen, Germany.
⁸¹ Department of Psychiatry, University of Pittsburgh, Pittsburgh, PA, USA.
⁸² Department of Human Genetics, University of Pittsburgh, Pittsburgh, PA, USA.
⁸³ Alzheimer's Disease Research Center, University of Pittsburgh, Pittsburgh, PA, USA.
⁸⁴ Institute of Genetics, Queen's Medical Centre, University of Nottingham, Nottingham, UK.
⁸⁵ Department of Neurology, Albert Einstein College of Medicine, Bronx, NY, USA.
⁸⁶ Section of Neuroscience, DIMEC-University of Parma, Parma, Italy.
⁸⁷ FERB-Alzheimer Center, Gazzaniga (Bergamo), Italy.
⁸⁸ Department of Pathology, University of Washington, Seattle, WA, USA.
⁸⁹ Elderly and Psychiatric Disorders Department, Medical University of Lodz, Lodz, Poland.
⁹⁰ Mercer's Institute for Research on Aging, St. James's Hospital and Trinity College, Dublin, Ireland.
⁹¹ St. James's Hospital and Trinity College, Dublin, Ireland.
⁹² Kaiser Permanente Washington Health Research Institute, Seattle, WA, USA.
⁹³ A.I. Virtanen Institute for Molecular Sciences, University of Eastern Finland, Kuopio, Finland.
⁹⁴ Departments of Medicine, Geriatrics, Gerontology and Neurology, University of Mississippi Medical Center, Jackson, MS, USA.
⁹⁵ Interdisciplinary Department of Medicine, Geriatric Medicine and Memory Unity, University of Bari, Bari, Italy.
⁹⁶ Department of Neurology, University of Washington, Seattle, WA, USA.
⁹⁷ Department of Geriatrics, Center for Aging Brain, University of Bari, Bari, Italy.
⁹⁸ Fundació per la Recerca Biomèdica i Social Mútua Terrassa, Terrassa, Barcelona, Spain.
⁹⁹ Memory Disorders Unit, Department of Neurology, Hospital Universitari Mutua de Terrassa, Terrassa, Barcelona, Spain.
¹⁰⁰ Department of Internal Medicine, Erasmus University Medical Center, Rotterdamt, the Netherlands.
¹⁰¹ Netherlands Consortium on Health Aging and National Genomics Initiative, Leiden, the Netherlands.
¹⁰² Department of Neurology, Mayo Clinic, Rochester, MN, USA.
¹⁰³ CHU Lille, Memory Center of Lille (Centre Mémoire de Ressources et de Recherche), Lille, France.
¹⁰⁴ Department of Clinical and Behavioral Neurology, Experimental Neuropsychobiology Laboratory, IRCCS Santa Lucia Foundation, Rome, Italy.
¹⁰⁵ School of Public Health, Human Genetics Center, University of Texas Health Science Center at Houston, Houston, TX, USA.
¹⁰⁶ Human Genome Sequencing Center, Baylor College of Medicine, Houston, TX, USA.
¹⁰⁷ Neurogenomics Division, Translational Genomics Research Institute, Phoenix, AZ, USA.
¹⁰⁸ Arizona Alzheimer's Consortium, Phoenix, AZ, USA.
¹⁰⁹ Banner Alzheimer's Institute, Phoenix, AZ, USA.
¹¹⁰ Department of Psychiatry, University of Arizona, Phoenix, AZ, USA.
¹¹¹ Institute for Translational Genomics and Population Sciences, Departments of Pediatrics and Medicine, Los Angeles BioMedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.
¹¹² Department of Clinical Sciences, University of Texas Southwestern Medical Center, Dallas, TX, USA.
¹¹³ University Paris Descartes, EA 4468, AP-HP, Geriatrics Department, Hôpital Broca, Paris, France.
¹¹⁴ Regional Neurogenetic Centre (CRN), ASP Catanzaro, Lamezia Terme, Italy.
¹¹⁵ Departments of Psychiatry, Medicine, Family & Community Medicine, South Texas Veterans Health Administration Geriatric Research Education & Clinical Center (GRECC), UT Health Science Center at San Antonio, San Antonio, TX, USA.
¹¹⁶ University of Bordeaux, Inserm 1219, Bordeaux, France.
¹¹⁷ Department of Neurology, Bordeaux University Hospital / CHU de Bordeaux, Bordeaux, France.
¹¹⁸ Department of Psychiatry, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
¹¹⁹ Inserm U1127, CNRS UMR 7225, Sorbonne Universités, UPMC Université Paris 06, UMRS 1127, Institut du Cerveau et de la Moelle Épinière, Paris, France.
¹²⁰ AP-HP, Department of Genetics, Pitié-Salpêtrière Hospital, Paris, France.
¹²¹ Section of Gerontology and Geriatrics, Department of Medicine, University of Perugia, Perugia, Italy.
¹²² Cambridge Institute for Medical Research, University of Cambridge, Cambridge, UK.
¹²³ Tanz Centre for Research in Neurodegenerative Disease, University of Toronto, Toronto, Ontario, Canada.
¹²⁴ Inserm U1061 Neuropsychiatry, La Colombière Hospital, Montpellier, France.
¹²⁵ Montpellier University, Montpellier, France.
¹²⁶ Department of Clinical Brain Sciences, University of Edinburgh, Edinburgh, UK.
¹²⁷ VA Puget Sound Health Care System/>GRECC, Seattle, WA, USA.
¹²⁸ Department of Psychiatry and Behavioral Sciences, University of Washington School of Medicine, Seattle, WA, USA.
¹²⁹ Institut de la Mémoire et de la Maladie d'Alzheimer and Institut du Cerveau et de la Moelle Épinière, Département de Neurologie, Hôpital de la Pitié-Salpêtrière, Paris, France.
¹³⁰ Institut des Neurosciences Translationnelles de Paris, Institut du Cerveau et de la Moelle Épinière, Paris, France.
¹³¹ Inserm, CNRS, UMR-S975, Institut du Cerveau et de la Moelle Epinière, Paris, France.
¹³² Sorbonne Universités, Université Pierre et Marie Curie, Hôpital de la Pitié-Salpêtrière, AP-HP, Paris, France.
¹³³ Ageing Group, Centre for Public Health, School of Medicine, Dentistry and Biomedical Sciences, Queen's University Belfast, Belfast, UK.
¹³⁴ Departments of Neurology, Pharmacology & Neuroscience, Texas Tech University Health Science Center, Lubbock, TX, USA.
¹³⁵ Department of Neurology, University of Michigan, Ann Arbor, MI, USA.
¹³⁶ Geriatric Research, Education and Clinical Center (GRECC), VA Ann Arbor Healthcare System (VAAAHS), Ann Arbor, MI, USA.
¹³⁷ Michigan Alzheimer Disease Center, Ann Arbor, MI, USA.
¹³⁸ Aging Research Center, Department of Neurobiology, Care Sciences and Society, Karolinska Institutet and Stockholm University, Stockholm, Sweden.
¹³⁹ Indiana Alzheimer's Disease Center, Indiana University School of Medicine, Indianapolis, IN, USA.
¹⁴⁰ Department of Medical and Molecular Genetics, Indiana University, Indianapolis, IN, USA.
¹⁴¹ Department of Neurology, Indiana University, Indianapolis, IN, USA.
¹⁴² Department of Radiology and Imaging Sciences, Indiana University, Indianapolis, IN, USA.
¹⁴³ Department of Psychiatry, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
¹⁴⁴ Geriatric Research, Education and Clinical Center (GRECC), University of Wisconsin, Madison, WI, USA.
¹⁴⁵ Department of Medicine, University of Wisconsin, Madison, WI, USA.
¹⁴⁶ Wisconsin Alzheimer's Disease Research Center, Madison, WI, USA.
¹⁴⁷ AXA Research Fund & Sorbonne University Chair, Paris, France.
¹⁴⁸ Sorbonne University, GRC n° 21, Alzheimer Precision Medicine (APM), AP-HP, Pitié-Salpêtrière Hospital, Paris, France.
¹⁴⁹ Brain & Spine Institute, Inserm U 1127, CNRS UMR 7225, Paris, France.
¹⁵⁰ Institute of Memory and Alzheimer's Disease, Department of Neurology, Pitié-Salpêtrière Hospital, AP-HP, Paris, France.
¹⁵¹ Grupo de Medicina Xenomica, Universidade de Santiago de Compostela, Centro Nacional de Genotipado, Centro de Investigación Biomédica en Red de Enfermedades Raras, Santiago de Compostela, Spain.
¹⁵² UK Dementia Research Institute, University of Cambridge, Cambridge, UK.
¹⁵³ Department of Neurological Sciences, Rush University Medical Center, Chicago, IL, USA.
¹⁵⁴ Department of Behavioral Sciences, Rush University Medical Center, Chicago, IL, USA.
¹⁵⁵ Rush Alzheimer's Disease Center, Rush University Medical Center, Chicago, IL, USA.
¹⁵⁶ Central Institute of Mental Health, Medical Faculty Mannheim, University of Heidelberg, Heidelberg, Germany.
¹⁵⁷ Civin Laboratory for Neuropathology, Banner Sun Health Research Institute, Phoenix, AZ, USA.
¹⁵⁸ Department of Neurology, University of Pittsburgh, Pittsburgh, PA, USA.
¹⁵⁹ Department of Psychology, University of Pittsburgh School of Medicine, Pittsburgh, PA, USA.
¹⁶⁰ Department of Pathology, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
¹⁶¹ Mesulam Center for Cognitive Neurology and Alzheimer's Disease, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
¹⁶² Genetic Epidemiology, QIMR Berghofer Medical Research Institute, Herston, Queensland, Australia.
¹⁶³ Swedish Medical Center, Seattle, WA, USA.
¹⁶⁴ Department of Neurology, University of California, San Francisco, San Francisco, CA, USA.
¹⁶⁵ Department of Neurology, Duke University, Durham, NC, USA.
¹⁶⁶ Departments of Biology, Brigham Young University, Provo, UT, USA.
¹⁶⁷ University of Kansas Alzheimer's Disease Center, University of Kansas Medical Center, Kansas City, KS, USA.
¹⁶⁸ Department of Experimental and Clinical Medicine, Neurological Institute, University of Pisa, Pisa, Italy.
¹⁶⁹ Department of Genetics and Genomic Sciences, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
¹⁷⁰ Department of Neuroscience, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
¹⁷¹ Department of Pathology and Immunology, Washington University, St. Louis, MO, USA.
¹⁷² Division of Psychiatry, University College London, London, UK.
¹⁷³ USF Health Byrd Alzheimer's Institute, University of South Florida, Tampa, FL, USA.
¹⁷⁴ Fred Hutchinson Cancer Research Center, Seattle, WA, USA.
¹⁷⁵ Department of Molecular Neuroscience, UCL, Institute of Neurology, London, UK.
¹⁷⁶ Mental Health & Behavioral Science Service, Bruce W. Carter VA Medical Center, Miami, FL, USA.
¹⁷⁷ Department of Neurodegenerative Disease, UCL Institute of Neurology, London, UK.
¹⁷⁸ Department of Neuroscience, Mayo Clinic, Jacksonville, FL, USA.
¹⁷⁹ Department of Neurology, University of Southern California, Los Angeles, CA, USA.
¹⁸⁰ Department of Neurology, Catholic University of Rome, Rome, Italy.
¹⁸¹ Department of Psychiatry and Behavioral Sciences, Miller School of Medicine, University of Miami, Miami, FL, USA.
¹⁸² Department of Neurology, University of California, Davis, Sacramento, CA, USA.
¹⁸³ Institute for Memory Impairments and Neurological Disorders, University of California, Irvine, Irvine, CA, USA.
¹⁸⁴ Wien Center for Alzheimer's Disease and Memory Disorders, Mount Sinai Medical Center, Miami Beach, FL, USA.
¹⁸⁵ Rush Institute for Healthy Aging, Department of Internal Medicine, Rush University Medical Center, Chicago, IL, USA.
¹⁸⁶ Department of Old Age Psychiatry, Institute of Psychiatry, Psychology and Neuroscience, King's College London, London, UK.
¹⁸⁷ Department of Primary Medical Care, University Medical Centre Hamburg-Eppendorf, Hamburg, Germany.
¹⁸⁸ Department of Pathology, University of Alabama at Birmingham, Birmingham, AL, USA.
¹⁸⁹ Sanders-Brown Center on Aging, Department of Biostatistics, University of Kentucky, Lexington, KY, USA.
¹⁹⁰ Department of Psychiatry, New York University, New York, NY, USA.
¹⁹¹ Department of Psychiatry and Psychotherapy, University of Cologne, Cologne, Germany.
¹⁹² Department of Psychiatry and Psychotherapy, Charité University Medicine, Berlin, Germany.
¹⁹³ Department of Neurosciences, University of California, San Diego, La Jolla, CA, USA.
¹⁹⁴ Department of Pathology and Laboratory Medicine, Emory University, Atlanta, GA, USA.
¹⁹⁵ Emory Alzheimer's Disease Center, Emory University, Atlanta, GA, USA.
¹⁹⁶ Department of Psychiatry, University of Freiburg, Freiburg, Germany.
¹⁹⁷ Neurogenetics Program, University of California, Los Angeles, Los Angeles, CA, USA.
¹⁹⁸ Department of Psychiatry, Washington University School of Medicine, St. Louis, MO, USA.
¹⁹⁹ Hope Center Program on Protein Aggregation and Neurodegeneration, Washington University School of Medicine, St. Louis, MO, USA.
²⁰⁰ Division of Genetics, Department of Medicine and Partners Center for Personalized Genetic Medicine, Brigham and Women's Hospital and Harvard Medical School, Boston, MA, USA.
²⁰¹ Department of Neurology, Washington University, St. Louis, MO, USA.
²⁰² Department of Genetics, Washington University, St. Louis, MO, USA.
²⁰³ Department of Neurology, Massachusetts General Hospital/Harvard Medical School, Boston, MA, USA.
²⁰⁴ Department of Pathology (Neuropathology), University of Pittsburgh, Pittsburgh, PA, USA.
²⁰⁵ Department of Neurology, University of Alabama at Birmingham, Birmingham, AL, USA.
²⁰⁶ Cologne Center for Genomics, University of Cologne, Cologne, Germany.
²⁰⁷ Department of Pathology, Duke University, Durham, NC, USA.
²⁰⁸ Department of Genome Sciences, University of Washington, Seattle, WA, USA.
²⁰⁹ Department of Medicine (Medical Genetics), University of Washington, Seattle, WA, USA.
²¹⁰ Sanders-Brown Center on Aging, College of Public Health, Department of Epidemiology, University of Kentucky, Lexington, KY, USA.
²¹¹ Unidad Clínica de Enfermedades Infecciosas y Microbiología, Hospital Universitario de Valme, Sevilla, Spain.
²¹² Department of Medicine (Biomedical Genetics), Boston University School of Medicine, Boston, MA, USA.
²¹³ Functional Genomics Center Zurich, ETH/University of Zurich, Zurich, Switzerland.
²¹⁴ Department of Neurology, Oregon Health &Science University, Portland, OR, USA.
²¹⁵ Department of Neurology, Portland Veterans Affairs Medical Center, Portland, OR, USA.
²¹⁶ Department of Pathology and Laboratory Medicine, University of California, Irvine, Irvine, CA, USA.
²¹⁷ Department of Pathology, Boston University School of Medicine, Boston University, Boston, MA, USA.
²¹⁸ Inserm U558, University of Toulouse, Toulouse, France.
²¹⁹ Department of Neuropsychology, University of California San Francisco, San Francisco, CA, USA.
²²⁰ Institute for Ageing and Health, Newcastle University, Newcastle upon Tyne, UK.
²²¹ Department of Neurobiology and Behavior, University of California, Irvine, Irvine, CA, USA.
²²² Department of Neurology, Emory University, Atlanta, GA, USA.
²²³ Institute for Stroke and Dementia Research, Klinikum der Universität München, Munich, Germany.
²²⁴ German Center for Neurodegenerative Diseases, Munich, Germany.
²²⁵ Cleveland Clinic Lou Ruvo Center for Brain Health, Cleveland Clinic, Cleveland, OH, USA.
²²⁶ Division of Neuroscience and Experimental Psychology, School of Biological Sciences, Faculty of Biology, Medicine and Health, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK.
²²⁷ Department of Pathology, University of Michigan, Ann Arbor, MI, USA.
²²⁸ Institute of Epidemiology, Helmholtz Zentrum München, German Research Center for Environmental Health, Neuherberg, Munich, Germany.
²²⁹ Helmholtz Center Munich, Institute of Epidemiology, Neuherberg, Munich, Germany.
²³⁰ Ludwig-Maximilians University Chair of Epidemiology, Munich, Germany.
²³¹ Joint Biobank Munich and KORA Biobank, Baltimore, MD, USA.
²³² Department of Psychiatry, Johns Hopkins University, Baltimore, MD, USA.
²³³ Human Genetics, Schools of Life Sciences and Medicine, University of Nottingham, Nottingham, UK.
²³⁴ Department of Medicine-Pulmonary, New York University, New York, NY, USA.
²³⁵ Department of Neurology, University of Miami, Miami, FL, USA.
²³⁶ Department of Pathology, University of California, San Diego, La Jolla, CA, USA.
²³⁷ Institute of Neurology, Catholic University of Sacred Hearth, Rome, Italy.
²³⁸ School of Nursing Northwest Research Group on Aging, University of Washington, Seattle, WA, USA.
²³⁹ Institute of Primary Care and Public Health, Cardiff University, University Hospital of Wales, Cardiff, UK.
²⁴⁰ Department of Neurology, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
²⁴¹ Cambridge Institute of Public Health, University of Cambridge School of Clinical Medicine, Cambridge, UK.
²⁴² Weill Institute for Neurosciences, Memory and Aging Center, University of California, San Francisco, San Francisco, CA, USA.
²⁴³ Institute of Social Medicine, Occupational Health and Public Health, University of Leipzig, Leipzig, Germany.
²⁴⁴ Department of Pathology, University of Southern California, Los Angeles, CA, USA.
²⁴⁵ Department of Pathology and Laboratory Medicine, University of California, Davis, Sacramento, CA, USA.
²⁴⁶ Institute of Psychiatry, Psychology and Neuroscienceó, King's College London, London, UK.
²⁴⁷ UK Dementia Research Institute, King's College London, London, UK.
²⁴⁸ Department of Psychiatry and Psychotherapy, University Medical Center Goettingen, Goettingen, Germany.
²⁴⁹ German Center for Neurodegenerative Diseases, Goettingen, Germany.
²⁵⁰ IBiMED, Medical Sciences Department, University of Aveiro, Aveiro, Portugal.
²⁵¹ Molecular Genetics Laboratory-Hospital, University of Central Asturias, Oviedo, Spain.
²⁵² Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, MN, USA.
²⁵³ Molecular Genetics Section, Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, MD, USA.
²⁵⁴ Department of Neurology, University of California, Irvine, Irvine, CA, USA.
²⁵⁵ Department of Neuroscience, Psychology, Drug Research and Child Health, University of Florence, Florence, Italy.
²⁵⁶ Centro di Ricerca, Trasferimento e Alta Formazione DENOTHE, University of Florence, Florence, Italy.
²⁵⁷ Department of Neurology, University of Colorado School of Medicine, Aurora, CO, USA.
²⁵⁸ IRCCS Fondazione Don Carlo Gnocchi, Florence, Italy.
²⁵⁹ Laboratory of Neuropsychiatry, IRCCS Santa Lucia Foundation, Rome, Italy.
²⁶⁰ Division of Neuropsychiatry, Department of Psychiatry and Behavioral Sciences, Baylor College of Medicine, Houston, TX, USA.
²⁶¹ Alzheimer's Disease Center, New York University, New York, NY, USA.
²⁶² Oxford Healthy Aging Project, Clinical Trial Service Unit, University of Oxford, Oxford, UK.
²⁶³ Department of Epidemiology, Columbia University, New York, NY, USA.
²⁶⁴ Oxford Project to Investigate Memory and Ageing, University of Oxford, Nuffield Department of Clinical Neurosciences, John Radcliffe Hospital, Oxford, UK.
²⁶⁵ Department of Neurology, Keck School of Medicine at the University of Southern California, Los Angeles, Los Angeles, CA, USA.
²⁶⁶ Department of Neurology, University of Texas Southwestern Medical Center, Dallas, TX, USA.
²⁶⁷ Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK.
²⁶⁸ Department of Pathology (Neuropathology), Rush University Medical Center, Chicago, IL, USA.
²⁶⁹ Department of Psychiatry, University of Southern California, Los Angeles, CA, USA.
²⁷⁰ Department of Pathology and Laboratory Medicine, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
²⁷¹ Department of Pathology, Johns Hopkins University, Baltimore, MD, USA.
²⁷² Sanders-Brown Center on Aging, Department of Neuroscience, University of Kentucky, Lexington, KY, USA.
²⁷³ Department of Neurology, University of California, Los Angeles, Los Angeles, CA, USA.
²⁷⁴ Department of Pathology and Laboratory Medicine, University of California, Los Angeles, Los Angeles, CA, USA.
²⁷⁵ Taub Institute on Alzheimer's Disease and the Aging Brain, Department of Pathology, Columbia University, New York, NY, USA.
²⁷⁶ Department of Psychiatry and Behavioral Sciences, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.
²⁷⁷ Department of Psychiatry and Behavioral Sciences, Duke University, Durham, NC, USA.
²⁷⁸ Department of Genetics, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.
²⁷⁹ Department of Human Genetics, Emory University, Atlanta, GA, USA.
²⁸⁰ Department of Pathology, Oregon Health & Science University, Portland, OR, USA.
²⁸¹ National Institute of Neurological Disorders and Stroke, Bethesda, MD, USA.
²⁸² Gottfried Schatz Research Center for Cell Signaling, Metabolism and Aging, Division of Molecular Biology and Biochemistry, Medical University Graz, Graz, Austria.
²⁸³ Gerontology and Geriatrics Research Laboratory, IRCCS Casa Sollievo della Sofferenza, San Giovanni Rotondo, Italy.
²⁸⁴ Department Geriatric Care, Orthogeriatrics and Rehabilitation, Galliera Hospital, Genova, Italy.
²⁸⁵ IdiPAZ, Instituto de Investigación Sanitaria la Paz, Madrid, Spain.
²⁸⁶ Centro de Biologia Molecular Severo Ochoa (CSIC-UAM), Madrid, Spain.
²⁸⁷ German Center for Neurodegenerative Diseases, Berlin, Germany.
²⁸⁸ Instituto di Ricovero e Cura a Carattere Scientifico, Associazione Oasi Maria Santissima Srl, Troina, Italy.
²⁸⁹ Center for Translational and Computational Neuroimmunology, Department of Neurology, Columbia University Medical Center, New York, NY, USA.
²⁹⁰ Memory Unit, Neurology Department and Sant Pau Biomedical Research Institute, Hospital de la Santa Creu i Sant Pau, Autonomous University Barcelona, Barcelona, Spain.
²⁹¹ Somerset Partnership NHS Trust, Somerset, UK.
²⁹² The Wellcome Trust Sanger Institute, Hinxton, Cambridge, UK.
²⁹³ Ronald M. Loeb Center for Alzheimer's Disease, Department of Neuroscience, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
²⁹⁴ University of Bristol Medical School, Learning & Research level 2, Southmead Hospital, Bristol, UK.
²⁹⁵ Department of Neurology, Mayo Clinic, Jacksonville, FL, USA.
²⁹⁶ Memory Research and Resources Center, CMRR de Bordeaux, Bordeaux, France.
²⁹⁷ Utah State University, Logan, UT, USA.
²⁹⁸ Department of Neurology, Erasmus MC University Medical Center, Rotterdam, the Netherlands.
²⁹⁹ Departments of Radiology, Erasmus MC University Medical Center, Rotterdam, the Netherlands.
³⁰⁰ Department of Research Rouvray Psychiatric Hospital, Sotteville-lès-Rouen, France.
³⁰¹ Department of Neurology, Medical University Graz, Graz, Austria.
³⁰² Center for Applied Genomics, Children's Hospital of Philadelphia, The Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA.
³⁰³ Division of Human Genetics, Department of Pediatrics, The Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA.
³⁰⁴ Department of Ophthalmology, Boston University School of Medicine, Boston University, Boston, MA, USA.
³⁰⁵ Department of Epidemiology, Boston University School of Public Health, Boston, MA, USA.
³⁰⁶ Department of Health Services, University of Washington, Seattle, WA, USA.
³⁰⁷ Kaiser Permanente, Washington Health Research Institute, Seattle, WA, USA.
³⁰⁸ Glenn Biggs Institute for Alzheimer's and Neurodegenerative Diseases, San Antonio, TX, USA.
³⁰⁹ Centre Hospitalier Universitaire de Lille, Lille, France.
³¹⁰ Inserm, U1167, RID-AGE-Risk Factors and Molecular Determinants of Aging-Related Diseases, Lille, France. jean-charles.lambert@pasteur-lille.fr.
³¹¹ Institut Pasteur de Lille, Lille, France. jean-charles.lambert@pasteur-lille.fr.
³¹² Univ. Lille, U1167-Excellence Laboratory LabEx DISTALZ, Lille, France. jean-charles.lambert@pasteur-lille.fr.
³¹³ John P. Hussman Institute for Human Genomics, University of Miami Miller School of Medicine, Miami, FL, USA. mpericak@miami.edu.

^# Contributed equally.

PMID: 31417202
PMCID: PMC7265117
DOI: 10.1038/s41588-019-0495-7

Abstract

An amendment to this paper has been published and can be accessed via a link at the top of the paper.

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Erratum for

Genetic meta-analysis of diagnosed Alzheimer's disease identifies new risk loci and implicates Aβ, tau, immunity and lipid processing.
Kunkle BW, Grenier-Boley B, Sims R, Bis JC, Damotte V, Naj AC, Boland A, Vronskaya M, van der Lee SJ, Amlie-Wolf A, Bellenguez C, Frizatti A, Chouraki V, Martin ER, Sleegers K, Badarinarayan N, Jakobsdottir J, Hamilton-Nelson KL, Moreno-Grau S, Olaso R, Raybould R, Chen Y, Kuzma AB, Hiltunen M, Morgan T, Ahmad S, Vardarajan BN, Epelbaum J, Hoffmann P, Boada M, Beecham GW, Garnier JG, Harold D, Fitzpatrick AL, Valladares O, Moutet ML, Gerrish A, Smith AV, Qu L, Bacq D, Denning N, Jian X, Zhao Y, Del Zompo M, Fox NC, Choi SH, Mateo I, Hughes JT, Adams HH, Malamon J, Sanchez-Garcia F, Patel Y, Brody JA, Dombroski BA, Naranjo MCD, Daniilidou M, Eiriksdottir G, Mukherjee S, Wallon D, Uphill J, Aspelund T, Cantwell LB, Garzia F, Galimberti D, Hofer E, Butkiewicz M, Fin B, Scarpini E, Sarnowski C, Bush WS, Meslage S, Kornhuber J, White CC, Song Y, Barber RC, Engelborghs S, Sordon S, Voijnovic D, Adams PM, Vandenberghe R, Mayhaus M, Cupples LA, Albert MS, De Deyn PP, Gu W, Himali JJ, Beekly D, Squassina A, Hartmann AM, Orellana A, Blacker D, Rodriguez-Rodriguez E, Lovestone S, Garcia ME, Doody RS, Munoz-Fernadez C, Sussams R, Lin H, Fairchild TJ, Benito YA, Holmes C, Karamujić-Čomić H, Fr… See abstract for full author list ➔ Kunkle BW, et al. Nat Genet. 2019 Mar;51(3):414-430. doi: 10.1038/s41588-019-0358-2. Epub 2019 Feb 28. Nat Genet. 2019. PMID: 30820047 Free PMC article.

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Author Correction: Genetic meta-analysis of diagnosed Alzheimer's disease identifies new risk loci and implicates Aβ, tau, immunity and lipid processing

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Author Correction: Genetic meta-analysis of diagnosed Alzheimer's disease identifies new risk loci and implicates Aβ, tau, immunity and lipid processing

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