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The Pseudomonas syringae Hrp pathogenicity island has a tripartite mosaic structure composed of a cluster of type III secretion genes bounded by exchangeable effector and conserved effector loci that contribute to parasitic fitness and pathogenicity in plants.
Alfano JR, Charkowski AO, Deng WL, Badel JL, Petnicki-Ocwieja T, van Dijk K, Collmer A. Alfano JR, et al. Proc Natl Acad Sci U S A. 2000 Apr 25;97(9):4856-61. doi: 10.1073/pnas.97.9.4856. Proc Natl Acad Sci U S A. 2000. PMID: 10781092 Free PMC article.